An Introduction to General Physiology With Practical Exercises

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This will be clear if we take a special case. Fats, as we saw, are esters in their chemical nature, and there are enzymes which bring about a splitting of esters in general into their component acids and alcohols. This they do by introducing water. Thus, taking ethyl acetate and putting Et for ethyl (C 2 H 5 ) and A for acetyl (CH 3 COO) we have :- Et A + HO Although this reaction only takes place rapidly under the action of a catalyst, it proceeds at a detectable rate if allowed to proceed by ...itself, but never completely. If we start with ethyl acetate and water in the proportions in which they combine, that is, if we take an equal multiple of the molecular weight of each, we find that after some days we can detect the presence of acetic acid (E. , p. 190). After a long time it will be found that no further change is going on, and that we have a state which we call equilibrium, . By estimating the amounts of the four com- 68 INTRODUCTION TO GENERAL PHYSIOLOGY poi>ents present, it is found that there are twice the number of molecules of the ester and water that there are of alcohol and acid.

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